Principal Component Analysis of diversity outcomes for 260 metacommunity simulations. Each “+” represents a simulated metacommunity. Simulations were ordinated by alpha-, beta-, and gamma-diversity (Jost 2007), and the environmental [E] and spatial [S] components of beta-diversity calculated using variation partitioning (Borcard et al. 2004, Peres-Neto et al. 2006). Simulation parameters that most influenced diversity outcomes are plotted as vectors. In the model, niche breadth (σtrait) is a parameter that determines the neutrality of the simulations.